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400  structures 8737  species 0  interactions 50487  sequences 628  architectures

Family: HSP70 (PF00012)

Summary: Hsp70 protein

Pfam includes annotations and additional family information from a range of different sources. These sources can be accessed via the tabs below.

This is the Wikipedia entry entitled "Hsp70". More...

This page is based on a Wikipedia article. The text is available under the Creative Commons Attribution/Share-Alike License.

This tab holds the annotation information that is stored in the Pfam database. As we move to using Wikipedia as our main source of annotation, the contents of this tab will be gradually replaced by the Wikipedia tab.

Hsp70 protein Provide feedback

Hsp70 chaperones help to fold many proteins. Hsp70 assisted folding involves repeated cycles of substrate binding and release. Hsp70 activity is ATP dependent. Hsp70 proteins are made up of two regions: the amino terminus is the ATPase domain and the carboxyl terminus is the substrate binding region.

Literature references

  1. Bukau B, Horwich AL; , Cell 1998;92:351-366.: The Hsp70 and Hsp60 chaperone machines. PUBMED:9476895 EPMC:9476895


Internal database links

External database links

This tab holds annotation information from the InterPro database.

InterPro entry IPR013126

Heat shock proteins, Hsp70 chaperones help to fold many proteins. Hsp70 assisted folding involves repeated cycles of substrate binding and release. Hsp70 activity is ATP dependent. Hsp70 proteins are made up of two regions: the amino terminus is the ATPase domain and the carboxyl terminus is the substrate binding region [ PUBMED:9476895 ].

Hsp70 proteins have an average molecular weight of 70kDa [ PUBMED:2686623 , PUBMED:2944601 , PUBMED:3282176 ]. In most species, there are many proteins that belong to the Hsp70 family. Some of these are only expressed under stress conditions (strictly inducible), while some are present in cells under normal growth conditions and are not heat-inducible (constitutive or cognate) [ PUBMED:2143562 , PUBMED:2841196 ]. Hsp70 proteins can be found in different cellular compartments (nuclear, cytosolic, mitochondrial, endoplasmic reticulum, for example).

This entry represents the Hsp70 family, and includes chaperone protein DnaK and luminal-binding proteins. It also includes heat shock protein 110 (Hsp110) from Caenorhabditis elegans which helps prevent the aggregation of denatured proteins in neurons [ PUBMED:19165329 ]. Also included is endoplasmic reticulum (ER) chaperone BiP (HSPA5) which is important for protein folding and quality control in the ER [ PUBMED:26655470 ].

Gene Ontology

The mapping between Pfam and Gene Ontology is provided by InterPro. If you use this data please cite InterPro.

Domain organisation

Below is a listing of the unique domain organisations or architectures in which this domain is found. More...

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Pfam Clan

This family is a member of clan Actin_ATPase (CL0108), which has the following description:

The actin-like ATPase domain forms an alpha/beta canonical fold. The domain can be subdivided into 1A, 1B, 2A and 2B subdomains. Subdomains 1A and 1B share the same RNAseH-like fold (a five-stranded beta-sheet decorated by a number of alpha-helices). Domains 1A and 2A are conserved in all members of this superfamily, whereas domain 1B and 2B have a variable structure and are even missing from some homologues [1]. Within the actin-like ATPase domain the ATP-binding site is highly conserved. The phosphate part of the ATP is bound in a cleft between subdomains 1A and 2A, whereas the adenosine moiety is bound to residues from domains 2A and 2B[1].

The clan contains the following 34 members:

Acetate_kinase Actin Actin_micro ALP_N AnmK BcrAD_BadFG Carbam_trans_N DDR DUF1464 DUF2229 EutA FGGY_C FGGY_N FtsA Fumble GDA1_CD39 Glucokinase Hexokinase_1 Hexokinase_2 HGD-D HSP70 Hydant_A_N Hydantoinase_A HypF_C MreB_Mbl MutL Pan_kinase PilM_2 Ppx-GppA RACo_C_ter ROK StbA T2SSL TsaD

Alignments

We store a range of different sequence alignments for families. As well as the seed alignment from which the family is built, we provide the full alignment, generated by searching the sequence database (reference proteomes) using the family HMM. We also generate alignments using four representative proteomes (RP) sets and the UniProtKB sequence database. More...

View options

We make a range of alignments for each Pfam-A family. You can see a description of each above. You can view these alignments in various ways but please note that some types of alignment are never generated while others may not be available for all families, most commonly because the alignments are too large to handle.

  Seed
(27)
Full
(50487)
Representative proteomes UniProt
(174649)
RP15
(9615)
RP35
(25503)
RP55
(45286)
RP75
(68314)
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PP/heatmap 1            

1Cannot generate PP/Heatmap alignments for seeds; no PP data available

Key: ✓ available, x not generated, not available.

Format an alignment

  Seed
(27)
Full
(50487)
Representative proteomes UniProt
(174649)
RP15
(9615)
RP35
(25503)
RP55
(45286)
RP75
(68314)
Alignment:
Format:
Order:
Sequence:
Gaps:
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Download options

We make all of our alignments available in Stockholm format. You can download them here as raw, plain text files or as gzip-compressed files.

  Seed
(27)
Full
(50487)
Representative proteomes UniProt
(174649)
RP15
(9615)
RP35
(25503)
RP55
(45286)
RP75
(68314)
Raw Stockholm Download   Download   Download   Download   Download      
Gzipped Download   Download   Download   Download   Download      

You can also download a FASTA format file containing the full-length sequences for all sequences in the full alignment.

HMM logo

HMM logos is one way of visualising profile HMMs. Logos provide a quick overview of the properties of an HMM in a graphical form. You can see a more detailed description of HMM logos and find out how you can interpret them here. More...

Trees

This page displays the phylogenetic tree for this family's seed alignment. We use FastTree to calculate neighbour join trees with a local bootstrap based on 100 resamples (shown next to the tree nodes). FastTree calculates approximately-maximum-likelihood phylogenetic trees from our seed alignment.

Note: You can also download the data file for the tree.

Curation and family details

This section shows the detailed information about the Pfam family. You can see the definitions of many of the terms in this section in the glossary and a fuller explanation of the scoring system that we use in the scores section of the help pages.

Curation View help on the curation process

Seed source: Prosite
Previous IDs: none
Type: Family
Sequence Ontology: SO:0100021
Author: Bateman A , Sonnhammer ELL
Number in seed: 27
Number in full: 50487
Average length of the domain: 404.40 aa
Average identity of full alignment: 29 %
Average coverage of the sequence by the domain: 80.60 %

HMM information View help on HMM parameters

HMM build commands:
build method: hmmbuild -o /dev/null HMM SEED
search method: hmmsearch -Z 57096847 -E 1000 --cpu 4 HMM pfamseq
Model details:
Parameter Sequence Domain
Gathering cut-off 20.6 20.6
Trusted cut-off 20.6 20.6
Noise cut-off 20.5 20.5
Model length: 599
Family (HMM) version: 22
Download: download the raw HMM for this family

Species distribution

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Archea Archea Eukaryota Eukaryota
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Viroids Viroids Unclassified sequence Unclassified sequence

Selections

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This visualisation provides a simple graphical representation of the distribution of this family across species. You can find the original interactive tree in the adjacent tab. More...

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Structures

For those sequences which have a structure in the Protein DataBank, we use the mapping between UniProt, PDB and Pfam coordinate systems from the PDBe group, to allow us to map Pfam domains onto UniProt sequences and three-dimensional protein structures. The table below shows the structures on which the HSP70 domain has been found. There are 400 instances of this domain found in the PDB. Note that there may be multiple copies of the domain in a single PDB structure, since many structures contain multiple copies of the same protein sequence.

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AlphaFold Structure Predictions

The list of proteins below match this family and have AlphaFold predicted structures. Click on the protein accession to view the predicted structure.

Protein Predicted structure External Information
A0A096R6Z8 View 3D Structure Click here
A0A0A0MPW9 View 3D Structure Click here
A0A0G2JVI3 View 3D Structure Click here
A0A0N7KD24 View 3D Structure Click here
A0A0N7KML3 View 3D Structure Click here
A0A0N7KSJ6 View 3D Structure Click here
A0A0P0V3A6 View 3D Structure Click here
A0A0P0V6T7 View 3D Structure Click here
A0A0P0W7F2 View 3D Structure Click here
A0A0P0WWL2 View 3D Structure Click here
A0A0P0WX25 View 3D Structure Click here
A0A0P0XZZ0 View 3D Structure Click here
A0A0P0YBH9 View 3D Structure Click here
A0A0R0EJN3 View 3D Structure Click here
A0A0R0FEK6 View 3D Structure Click here
A0A0R0FH00 View 3D Structure Click here
A0A0R0FY49 View 3D Structure Click here
A0A0R0G7C6 View 3D Structure Click here
A0A0R0G7H5 View 3D Structure Click here
A0A0R0G7H5 View 3D Structure Click here
A0A0R0GDY9 View 3D Structure Click here
A0A0R0GRW1 View 3D Structure Click here
A0A0R0GRW1 View 3D Structure Click here
A0A0R0H2C6 View 3D Structure Click here
A0A0R0HS21 View 3D Structure Click here
A0A0R0HS21 View 3D Structure Click here
A0A0R0HU88 View 3D Structure Click here
A0A0R0HU88 View 3D Structure Click here
A0A0R0I1Z7 View 3D Structure Click here
A0A0R0I1Z7 View 3D Structure Click here
A0A0R0JEY3 View 3D Structure Click here
A0A0R0KKU3 View 3D Structure Click here
A0A0R0KUH0 View 3D Structure Click here
A0A0R0KUH0 View 3D Structure Click here
A0A0R0KUH0 View 3D Structure Click here
A0A0R4J4D6 View 3D Structure Click here
A0A0R4J626 View 3D Structure Click here
A0A1D6F848 View 3D Structure Click here
A0A1D6F848 View 3D Structure Click here
A0A1D6F848 View 3D Structure Click here
A0A1D6FM90 View 3D Structure Click here
A0A1D6FN98 View 3D Structure Click here
A0A1D6FNB8 View 3D Structure Click here
A0A1D6GK64 View 3D Structure Click here
A0A1D6GMN4 View 3D Structure Click here
A0A1D6GSN4 View 3D Structure Click here
A0A1D6HHX7 View 3D Structure Click here
A0A1D6HTL5 View 3D Structure Click here
A0A1D6HTR0 View 3D Structure Click here
A0A1D6HUR4 View 3D Structure Click here
A0A1D6I1V7 View 3D Structure Click here
A0A1D6I2R4 View 3D Structure Click here
A0A1D6I793 View 3D Structure Click here
A0A1D6I852 View 3D Structure Click here
A0A1D6J1L4 View 3D Structure Click here
A0A1D6J4I6 View 3D Structure Click here
A0A1D6J7R6 View 3D Structure Click here
A0A1D6J7R7 View 3D Structure Click here
A0A1D6J7S4 View 3D Structure Click here
A0A1D6J7S6 View 3D Structure Click here
A0A1D6J7S6 View 3D Structure Click here
A0A1D6JY78 View 3D Structure Click here
A0A1D6KE29 View 3D Structure Click here
A0A1D6LPI6 View 3D Structure Click here
A0A1D6LZY7 View 3D Structure Click here
A0A1D6M007 View 3D Structure Click here
A0A1D6MUE8 View 3D Structure Click here
A0A1D6MUP2 View 3D Structure Click here
A0A1D6N7I4 View 3D Structure Click here
A0A1D6PAX7 View 3D Structure Click here
A0A1D6PAX7 View 3D Structure Click here
A0A1D6PKR9 View 3D Structure Click here
A0A1D6Q8C9 View 3D Structure Click here
A0A1D8PG96 View 3D Structure Click here
A0A1D8PGU0 View 3D Structure Click here
A0A1D8PRH1 View 3D Structure Click here
A0A2R8QHY3 View 3D Structure Click here
A0A2R8QJR3 View 3D Structure Click here
A0A368UHA7 View 3D Structure Click here
A0A368UHA7 View 3D Structure Click here
A0A368UHA7 View 3D Structure Click here
A0A368UHI2 View 3D Structure Click here
A0A368UHI2 View 3D Structure Click here
A4HRI7 View 3D Structure Click here
A4HRI7 View 3D Structure Click here
A4I219 View 3D Structure Click here
A4I219 View 3D Structure Click here
A4I253 View 3D Structure Click here
A4I3J9 View 3D Structure Click here
A4I412 View 3D Structure Click here
A4I417 View 3D Structure Click here
A4I4I1 View 3D Structure Click here
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A4I5S9 View 3D Structure Click here
A4I5T0 View 3D Structure Click here
A4IC10 View 3D Structure Click here
A5PMG2 View 3D Structure Click here
B0S610 View 3D Structure Click here
B0UXR9 View 3D Structure Click here
B3H683 View 3D Structure Click here
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B6SZ69 View 3D Structure Click here
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B9G4B3 View 3D Structure Click here
C0H5H0 View 3D Structure Click here
C4J410 View 3D Structure Click here
C4JBB8 View 3D Structure Click here
E9AGQ5 View 3D Structure Click here
E9AHH0 View 3D Structure Click here
E9AHH1 View 3D Structure Click here
F1LZI1 View 3D Structure Click here
F1QFC0 View 3D Structure Click here
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F4HQD4 View 3D Structure Click here
F4JMJ1 View 3D Structure Click here
G5EC99 View 3D Structure Click here
G5ECU5 View 3D Structure Click here
I1JAU2 View 3D Structure Click here
I1JGR5 View 3D Structure Click here
I1K668 View 3D Structure Click here
I1K670 View 3D Structure Click here
I1KG78 View 3D Structure Click here
I1KL72 View 3D Structure Click here
I1KPN3 View 3D Structure Click here
I1KPN5 View 3D Structure Click here
I1KQV7 View 3D Structure Click here
I1KV87 View 3D Structure Click here
I1KY70 View 3D Structure Click here
I1LQR4 View 3D Structure Click here
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I1ME25 View 3D Structure Click here
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I1MT10 View 3D Structure Click here
I1N0Q9 View 3D Structure Click here
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K7K753 View 3D Structure Click here
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K7L2A0 View 3D Structure Click here
K7LZH1 View 3D Structure Click here
K7MQ03 View 3D Structure Click here
K7MQ03 View 3D Structure Click here
K7MQ03 View 3D Structure Click here
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K7VW90 View 3D Structure Click here
M0R8M9 View 3D Structure Click here
M0RCB1 View 3D Structure Click here
M9MSL3 View 3D Structure Click here
O07239 View 3D Structure Click here
O24581 View 3D Structure Click here
O35162 View 3D Structure Click here
O35162 View 3D Structure Click here
O45246 View 3D Structure Click here
O46067 View 3D Structure Click here
O53538 View 3D Structure Click here
O59838 View 3D Structure Click here
O59855 View 3D Structure Click here
O65719 View 3D Structure Click here
O88600 View 3D Structure Click here
O88600 View 3D Structure Click here
O95757 View 3D Structure Click here
O95757 View 3D Structure Click here
O97125 View 3D Structure Click here
P02825 View 3D Structure Click here
P06761 View 3D Structure Click here
P09435 View 3D Structure Click here
P09446 View 3D Structure Click here
P0A6Y8 View 3D Structure Click here
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P0CS90 View 3D Structure Click here
P0DMV8 View 3D Structure Click here
P0DMV9 View 3D Structure Click here
P0DMW0 View 3D Structure Click here
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P10591 View 3D Structure Click here
P10592 View 3D Structure Click here
P11021 View 3D Structure Click here
P11141 View 3D Structure Click here
P11142 View 3D Structure Click here
P11146 View 3D Structure Click here
P11147 View 3D Structure Click here
P11484 View 3D Structure Click here
P14659 View 3D Structure Click here
P16474 View 3D Structure Click here
P16627 View 3D Structure Click here
P17066 View 3D Structure Click here
P17156 View 3D Structure Click here
P17879 View 3D Structure Click here
P20029 View 3D Structure Click here
P20163 View 3D Structure Click here
P22202 View 3D Structure Click here
P22774 View 3D Structure Click here
P22953 View 3D Structure Click here
P22954 View 3D Structure Click here
P24067 View 3D Structure Click here
P26413 View 3D Structure Click here
P27420 View 3D Structure Click here
P29843 View 3D Structure Click here
P29844 View 3D Structure Click here
P29845 View 3D Structure Click here
P32589 View 3D Structure Click here
P32590 View 3D Structure Click here
P34931 View 3D Structure Click here
P34932 View 3D Structure Click here
P34932 View 3D Structure Click here
P36016 View 3D Structure Click here
P36415 View 3D Structure Click here
P36604 View 3D Structure Click here
P36928 View 3D Structure Click here
P36928 View 3D Structure Click here
P38646 View 3D Structure Click here
P38647 View 3D Structure Click here
P38788 View 3D Structure Click here
P39987 View 3D Structure Click here
P40150 View 3D Structure Click here
P41797 View 3D Structure Click here
P46587 View 3D Structure Click here
P48721 View 3D Structure Click here
P48722 View 3D Structure Click here
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P48723 View 3D Structure Click here
P48741 View 3D Structure Click here
P54652 View 3D Structure Click here
P55063 View 3D Structure Click here
P63017 View 3D Structure Click here
P63018 View 3D Structure Click here
P77319 View 3D Structure Click here
P82910 View 3D Structure Click here
P83784 View 3D Structure Click here
P87142 View 3D Structure Click here
P9WMJ9 View 3D Structure Click here
P9WMJ9 View 3D Structure Click here
Q05036 View 3D Structure Click here
Q05931 View 3D Structure Click here
Q0DJB9 View 3D Structure Click here
Q0IU43 View 3D Structure Click here
Q0VDF9 View 3D Structure Click here
Q10061 View 3D Structure Click here
Q10265 View 3D Structure Click here
Q10284 View 3D Structure Click here
Q10NA1 View 3D Structure Click here
Q10NA5 View 3D Structure Click here
Q10NA9 View 3D Structure Click here
Q10PW8 View 3D Structure Click here
Q10SR3 View 3D Structure Click here
Q1MT77 View 3D Structure Click here
Q20752 View 3D Structure Click here
Q22515 View 3D Structure Click here
Q22758 View 3D Structure Click here
Q2FXZ2 View 3D Structure Click here
Q2FXZ2 View 3D Structure Click here
Q2QV45 View 3D Structure Click here
Q2R9K4 View 3D Structure Click here
Q39043 View 3D Structure Click here
Q4CK86 View 3D Structure Click here
Q4CL09 View 3D Structure Click here
Q4CPR7 View 3D Structure Click here
Q4CS27 View 3D Structure Click here
Q4CU95 View 3D Structure Click here
Q4CUA0 View 3D Structure Click here
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Q4D118 View 3D Structure Click here
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Q4D673 View 3D Structure Click here
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Q4DGL0 View 3D Structure Click here
Q4DI67 View 3D Structure Click here
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Q4DXG9 View 3D Structure Click here
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Q4E175 View 3D Structure Click here
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Q53NM9 View 3D Structure Click here
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Q54BD8 View 3D Structure Click here
Q54BE0 View 3D Structure Click here
Q54BE1 View 3D Structure Click here
Q54MR6 View 3D Structure Click here
Q54TS8 View 3D Structure Click here
Q556U6 View 3D Structure Click here
Q557E0 View 3D Structure Click here
Q5A397 View 3D Structure Click here
Q5A678 View 3D Structure Click here
Q5KQJ9 View 3D Structure Click here
Q5RGE6 View 3D Structure Click here
Q5RJ08 View 3D Structure Click here
Q61316 View 3D Structure Click here
Q61316 View 3D Structure Click here
Q61696 View 3D Structure Click here
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Q63617 View 3D Structure Click here
Q66HA8 View 3D Structure Click here
Q69TV0 View 3D Structure Click here
Q6AYB4 View 3D Structure Click here
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Q6L509 View 3D Structure Click here
Q6L590 View 3D Structure Click here
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Q6P3L3 View 3D Structure Click here
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Q6PH56 View 3D Structure Click here
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Q6Z058 View 3D Structure Click here
Q6Z7B0 View 3D Structure Click here
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Q6ZFX8 View 3D Structure Click here
Q75HQ0 View 3D Structure Click here
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Q7ZUW2 View 3D Structure Click here
Q84TA1 View 3D Structure Click here
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Q8IB24 View 3D Structure Click here
Q8IC01 View 3D Structure Click here
Q8IC01 View 3D Structure Click here
Q8II24 View 3D Structure Click here
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Q8T869 View 3D Structure Click here
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Q92598 View 3D Structure Click here
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